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The existence of genetic correlations can be established via quantitative genetic breeding designs or by way of correlated phenotypic responses to selection. This suggests that lifespan and early reproduction are genetically negatively coupled, e.g. through antagonistic pleiotropic alleles (e.g., Flatt 2011, Fabian and Flatt 2011). But how can we reconcile the fact that VA is massive whilst at the very same time h2 is compact? A probably explanation for the low heritability of life history traits is that, although VA (the numerator) is substantial, VP (the denominator) is a lot larger than VA. A female North Pacific Giant Octopus (Enteroctopus dofleini) lives three here to four years it lays thousands of eggs in a single bout and then dies. By contrast, a mature Coast Redwood Tree (Sequoia sempervirens) lives for lots of hundreds of years and produces millions of seeds each year (Figure 1). As these two examples illustrate, organisms differ considerably in how they create, the time they take to develop, when they become mature, how numerous offspring of a specific size they generate, and how lengthy they live. With each other, the age-, size-, or stage-specific patterns of development, growth, maturation, reproduction, survival, and lifespan define an organism's life cycle, its life history.

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